Kanehiro Kitayama., Masayuki Ushio and Shin-ichiro Aiba (2020) Temperature is a dominant driver of distinct annual seasonality of leaf litter production of equatorial tropical rain forests. Journal of Ecology, 2020;00:1-10.

Hatt S., Francis F., Xu Q., Wang S., Osawa N. (2020) Perennial flowering strips for conservation biological control of insect pests: from picking and mixing flowers to tailored functional diversity. In: Gao Y., Hokkanen H., Menzler-Hokkanen I. (Eds), Integrative Biological Control, Progress in Biological Control Series, Springer, 57-71.

Xu Q., Wang S., Li S., Hatt S. (2020) Conservation biological control in organic greenhouse vegetables. In: Gao Y., Hokkanen H., Menzler-Hokkanen I. (Eds), Integrative Biological Control, Progress in Biological Control Series, Springer, 133-144.

Li S., Jaworski C. C., Hatt S., Zhang F., Desneux N., Wang S. (2020). Flower strips adjacent to greenhouses help reduce pest populations and insecticide applications inside organic commercial greenhouses. Journal of Pest Science.

Sakata Y, Kobayashi K, Makita A (2020) Insect assemblages on flowering patches of 12 bamboo species. Journal of Asia-Pacific Entomology (in press)



Bayoumy M. H., Osawa N.& Hatt S. (2019) Fitness costs of reflex bleeding in the ladybird beetle Harmonia axyridis: the role of paternal effects. Insect Science, 00, 1-14

Kobayashi K, Kobayashi T. (2019) Extremely high density of flowered culms observed for a moso bamboo (Phyllostachys edulis) stand in Tosa city, Kochi prefecture, western Japan. Applied Forest Science 28 : 11-15

Amada G, Kobayashi K, Izuno A, Mukai M, Ostertag R, Kitayama K, & Onoda Y(2019). Leaf trichomes in Metrosideros polymorpha can contribute to avoiding extra water stress by impeding gall formations. Annals of Botany, 125(3), 533-542

Hatt S., Osawa N. (2019). Beyond ‘‘greening’’: which paradigms shape sustainable pest management strategies in the European Union? BioControl, 64(4), 343-355.

Hatt S., Osawa N. (2019). The role of Perilla frutescens flowers on fitness traits of the ladybird beetle Harmonia axyridis. BioControl, 64(4), 381-390.

Hatt S., Xu Q., Francis F., Osawa N. (2019). Aromatic plants of East Asia to enhance natural enemies towards biological control of insect pests. A review. Entomologia Generalis, 38(4), 275-315.

Hatt S., Uyttenbroeck R., Lopes T., Mouchon P., Osawa N., Piqueray J., Monty A., Francis F. (2019). Identification of flower functional traits affecting abundance of generalist predators in perennial multiple species wildflower strips. Arthropod-Plant Interactions, 13, 127-137.

Mana Mukai, Yoshimi Sawada, Kanehiro Kitayama & Shin-ichiro Aiba (2019). Productivity and morphological traits of fine roots in forest ecosystems along an elevation gradient of Yakushima Island. Journal of Forest Research, 24(1), 35-41



Hiroki Ito, Naoya Osawa (2018) A field study of the colony composition of the wood-feeding cockroach Panesthia angustipennis spadica (Blattodea: Blaberidae).Applied Entomology and Zoology.

Kitayama Kanehiro, Fujiki Shogoro, Aoyagi Ryota, Imai Nobuo, Sugau John, Titin Jupiri, Nilus Reuben, Lagan Peter, Sawada Yoshimi, Ong Robert, Kugan Frederick, Mannan Sam (2018) Biodiversity Observation for Land and Ecosystem Health (BOLEH): A Robust Method to Evaluate the Management Impacts on the Bundle of Carbon and Biodiversity Ecosystem Services in Tropical Production Forests, Sustainability, 10(11)

Yuki Tsujii, Kanehiro Kitayama (2018) Relationships of phosphorus concentration in reproductive organs with soil phosphorus availability for tropical rain-forest trees on Mount Kinabaru, Borneo. Journal of Tropical Ecology.

Kobayashi K, Kitayama K, Onoda Y (2018) A simple method to estimate the rate of the bamboo expansion based on one-time measurement of spatial distribution of culms. Ecological Research.

Agus S Jati, Samejima H, Fujiki S, Kurniawan Y, Aoyagi R and Kitayama K (2018) Effect of logging on wildlife communities in certified tropical rainforests in East Kalimantan, Indonesia. Forest Ecology and Management.

Fujiki S, Nishio S, Kei-ichi Okada, Nais J, Repin R, Kitayama K (2018) Estimation of the Spatiotemporal Patterns of Vegetation and Associated Ecosystem Services in a Bornean Montane Zone Using Three Shifting-Cultivation Scenarios. Land



Mori T., Imai N., Kitayama K. (2018) A preliminary report: does reduced impact logging (RIL) mitigate non-CO2 greenhouse gas emissions from natural production forests? Tropics 27:25-31.

Mori T., Imai N., Yokoyama D., Mukai M., Kitayama K. (2017) Effects of selective logging and application of phosphorus and nitrogen of C02, CH4 and N2O in lowland tropical rainforests of Borneo. Journal of Tropical Forest Science 29: 248-256.

Yokoyama D, Mori T, Wagai R, Hiradate S, Kitayama K (2017) Characteristics of phosphorus fractions in the soils derived from sedimentary and serpentinite rocks in lowland tropical rain forests, Borneo. Soil Science and Plant Nutrition.

Tsujii Y, Onoda Y, Kitayama K. (2017). Phosphorus and nitrogen resorption from different chemical fractions in senescing leaves of tropical tree species on Mount Kinabalu, Borneo. Oecologia.

Aoyagi R, Imai N, Fujiki S, John B. Sugau, Joan T. Pereira,Kitayama K (2017) The mixing ratio of tree functional groups as a new index for biodiversity monitoring in Bornean production forests. Forest Ecology and Management.

Tsujii Y, Oikawa M & Kitayama K (2017) Significance of the localization of phosphorus among tissues on a cross-section of leaf lamina of Bornean tree species for phosphorus-use efficiency. Journal of Tropical Ecology.

Amada G, Onoda Y, Ichie T & Kitayama K (2017)Influence of leaf trichomes on boundary layer conductance and gas-exchange characteristics in Metrosideros polymorpha (Myrtaceae). Biotropica

Endo N., Osawa N. (2017) Asymmetric song responses in coexisting Japanese cicadas. Bioacourstics 27.

Fujiki S., Shogo Nishio, Kei-ichi Okada, Jamili Nais and Kanehiro Kitayama (2017). Plant communities and ecosystem processes in a succession-altitude matrix after shifting cultivation in the tropical montane forest zone of northern Borneo. Journal of Tropical Ecology doi:10.1017/S0266467416000547

Ito H., Osawa N. (2017) The effects of aggregation on survival and growth rate in the wood-feeding cockroach Panesthia angustipennis spadica (Order: Blaberidae). Entomological Science 20 (in press).

Izuno A, Kitayama K, Onoda Y, Tsujii Y, Hatakeyama M, Nagano AJ, Honjo MN, Shimizu-Inatsugi R, Kudoh H, Shimizu KK, Isagi Y (2017) The population genomic signature of environmental selection with gene flow in an ecologically divergent tree species Metrosideros polymorpha (Myrtaceae). Molecular Ecology (in press).

Okada K., Shin-ichro Aiba, Kanehiro Kitayama (2017). Influence of temperature and soil nitrogen and phosphorus availabilities on fine-root productivity in tropical rainforests on Mount Kinabalu, Borneo. Ecological Research, doi:10.1007/s11284-016-1425-0
Onoda Y, Wright IJ, Evans JR, Hikosaka K, Kitajima K, Niinemets U, Poorter H, Tosens T, Westoby M. (2017) Physiological and structural tradeoffs underlying the leaf economics spectrum. New Phytologist

Sun L., Mioko Ataka, Yuji Kominami, Kenichi Yoshimura. (2017). Relationship between fine-root exudation and respiration of two Quercus species in a Japanese temperate forest. Tree Physiol doi: 10.1093/treephys/tpx026

Sun L., Yuji Kominami, Kenichi Yoshimura, Kanehiro Kitayama. (2017). Root-exudate flux variations among four co-existing canopy species in a temperate forest, Japan. Ecological Research 32: 331-339 doi:10.1007/s11284-017-1440-9

Ueda M, Onoda Y, Kamiyama C & Hikosaka K (2017) Decades-long effects of high CO2 concentration on soil nitrogen dynamics at a natural CO2 spring. Ecological Research (in press)

Ushio M., Aiba S., Takeuchi Y., Iida Y., Matsuoka S., Repin R., Kitayama K. (2017) Plant-soil feedbacks and the dominance of conifers in a tropical montane forest in Borneo. Ecological Monographs 87: 105-129.

Yokoyama D., Nobuo Imai, Kanehiro Kitayama. (2017). Effects of nitrogen and phosphorus fertilization on the activities of four different classes of fine-root and soil phosphatases in Bornean tropical rain forests. Plant and soil


Aiba M, Kurokawa H, Onoda Y, Oguro M, Nakashizuka T, Masaki T (2016) Context-dependent changes in the functional composition of tree communities along successional gradients after land-use change. Journal of Ecology 104: 1347?1356
Aoyagi R., Kitayama K. (2016) Nutrient allocation among plant organs across 13 tree species in three Bornean rain forests with contrasting nutrient availabilities. Journal of Plant Research 129: 675-684.

Aoyagi R., Nobuo Imai, Tatsuyuki Seino, Kanehiro Kitayama (2016), Soil nutrients and size-dependent tree dynamics of tropical lowland forests on volcanic and sedimentary substrates in Sabah, Malaysian Borneo, Tropics, 25, pp43-52.

Fujiki S., Aoyagi R., Tanaka A., Imai N., Kusma A.D., Kurniawan Y., Lee Y.F, Sugau J.B., Pereira, J.T., Samejima H., Kitayama K. (2016) Large-scale mapping of tree-community composition as a surrogate of forest degradation in Bornean tropical rain forests. Land.

Fujiki S., Okada K., Nishio S., Kitayama K. (2016) Estimation of the stand ages of tropical secondary forests after shifting cultivation based on the combination of WorldView-2 and time-series Landsat images. ISPRS Journal of Photogrammetry and Remote Sensing 119: 280-293.

Iidzuka H., Goto H., Osawa N. (2016) Gallery diameter of ambrosia beetles (Coleoptera: Scolytidae, Platypodidae) and insect fauna in Quercus serrata suffering from Japanese oak wilt. Applied Entomology and Zoology 51: 421-427.

Iidzuka H., Goto H., Yamasaki M., Osawa N. (2016) Wood boring beetles (Coleoptera: Scolytidae, Platypodidae) captured in ethanol-baited traps at the natural forest in Japan. Applied Entomology and Zoology 51: 347-352.
Iidzuka H., Osawa N. (2016) Horizontal niche differentiation of ambrosia beetles (Coleoptera: Scolytidae and Platypodidae) within the standing trunk of Quercus serrata. Journal of Forest Research 21: 319-323.

Imai N., Samejima H., Malcom S.,Tanaka A., Sugau J., Pereira J., Kitayama K. (2016) Response of tree species diversity to disturbance in humid tropical forests of Borneo. Journal of Vegetation Science 27: 739?748. doi:10.1111/jvs.12401

Lin C.-P., Osawa N., Kitano T., Huang S.-C. (2016) Coexistence of two species of Harmonia ladybird beetles (Coleoptera: Coccinellidae) on the Taiwan Red Pine Pinus taiwanensis in Alishan of Taiwan: rediscovery of Harmonia yedoensis and a new record of Harmonia quadripunctata. Formosan Entomologist 36: 1-5.

Mori T., Yokoyama D., Kitayama K. (2016) Contrasting effects of exogenous phosphorus application on N2O emissions from two tropical forest soils with contrasting phosphorus availability. SpringerPlus5: 1237 DOI 10.1186/s40064-016-2587-5

Mori T., Wachrinrat C, Staporn D, Meunpong P, Suebsai W, Boonsri K, and Kitayama K. (2016). Seasonal changes in soil respiration and microbial biomass in five tropical tree plantations in Thailand. Topics, 25(2): 85−89.

Mukai M., Aiba S., Kitayama K. (2016) Soil-nutrient availability and the nutrient-use efficiencies of the forests along an altitudinal gradient on Yakushima Island. Ecological Research
Noriyuki S., Osawa N. (2016) Reproductive interference and niche partitioning in aphidophagous insects. Psyche 216: Article ID 4751280, 15 pages.

Roy H. E., Brown P. M. J., Adriaens T., ...Osawa N., ....(2016) The harlequin ladybird, Harmonia axyridis: global perspectives on invasion history and ecology. Biological Invasions 18: 997-1044.

Sullivan M.J.P. ...Kitayama K., .... (2016) Diversity and carbon storage across the tropical forest biome. Scientific Reports 6:39102 | DOI: 10.1038/srep39102

Tatsuno M., Osawa N. (2016) Flower visitation patterns of the coexisting honey bees Apis cerana japonica and Apis mellifera (Hymenoptera: Apidae). Entomological Science 19: 255-267.

Toosi M., Rasekh A., Osawa N. (2016) Intraguild predation by Hippodamia variegata (Col.: Coccinellidae) on the parasitoid wasp, Lysiphlebus fabarum (Hym.: Braconidae). Journal of Entomological Society of Iran 36: 205-216.

Tsujii Y., Yusuke Onoda, Ayako Izuno, Yuji Isagi, Kanehiro Kitayama(2016) A quantitative analysis of phenotypic variations of Metrosideros polymorpha within and across populations along environmental gradients on Mauna Loa, Hawaii. Oecologia 180:1049?1059.

de los Santos CB, Onoda Y, Vergara JJ, Perez-Llorens JL, Bouma TJ, La Nafie YA, Cambridge ML, Brun FG (2016) A comprehensive analysis of mechanical and morphological traits in one third of world seagrass species. Marine Ecology Progress Series 551:81-94 LINK

Aiba S., Sawada Y., Takyu M., Seino T., Kitayama K., Repin R. (2015) Structure, floristics and diversity of tropical montane rain forests over ultramafic soils on Mount Kinabalu (Borneo) compared with those on non-ultramafic soils. Australian Journal of Botany 63: 191?203

Aoyagi R., Kanehiro Kitayama (2015), Traits associated with nutrient impoverishment and shade-tolerance in tree juveniles of three Bornean rain forests with contrasting nutrient availability. Journal of Tropical Ecology 31, pp231-241,.
Hidaka A., Kitayama K. (2015) Physiological linkage in co-variation of foliar nitrogen and phosphorus in tropical tree species along a gradient of soil phosphorus availability. Journal of Tropical Ecology 31: 221-229

Ishihara MI, Utsugi H, Tanouchi H, Aiba M, Kurokawa H, Onoda Y, Nagano M, Umehara T, Ando M, Miyata R, Hiura, T. (2015) Efficacy of generic allometric equations for estimating biomass: a test in Japanese natural forests. Ecological Applications 25:1433?1446.

Kitayama K., Tsujii Y., Aoyagi R., Aiba S. (2015) Long-term C, N and P allocation to reproduction in Bornean tropical rain forests. Journal of Ecology 103: 606-615.

Kunstler G, Falster D, Coomes DA, Hui F, Kooyman RM, Laughlin DC, Poorter L, Vanderwel M, Vieilledent G, Wright SJ, Aiba M, Baraloto C, Caspersen J, Cornelissen JHC, Gourlet-Fleury S, Hanewinkel M, Herault B, Kattge J, Kurokawa H, Onoda Y, Penuelas J, Poorter H, Uriarte M, Richardson S, Ruiz-Benito P, Sun IF, Stahl G, Swenson NG, Thompson J, Westerlund B, Wirth C, Zavala MA, Zeng H, Zimmerman JK, Zimmermann NE, Westoby M. (2015) Plant functional traits have globally consistent effects on competition. Nature. 529, 204?207

Langner A.,Achard F., Vancutsem C., Pekel J.-F.,Simonetti D., Grassi G., Kitayama K., Nakayama M. (2015) Assessment of Above-Ground Biomass of Borneo Forests through a New Data-Fusion Approach Combining Two Pan-Tropical Biomass Maps. Land 4: 656-669; doi:10.3390/land4030656

Lin ...Onoda Y.,...(2015) Optimal stomatal behaviour around the world. Nature Climate Change 5, 459?464.
Noriyuki S., Osawa N. (2015) Geographic variation of color polymorphism in the two sibling ladybird species, Harmonia yedoensis and Harmonia axyridis (Coleoptera: Coccinellidae). Entomological Science 18: 502-508.

Onoda Y, Schieving F & Anten NPR. (2015) A novel method of measuring leaf epidermis and mesophyll stiffness shows the ubiquitous nature of the sandwich structure of leaf laminas in broad-leaved angiosperm species. Journal of Experimental Botany 66: 2487?2499. プレスリリース記事 JSTサイエンスポータル 測定手法の説明 新聞電子版(朝日新聞、財経新聞、京都新聞、日経新聞)

Osawa N., Kagami N., Noriyuki N., Tuno N. (2015) Size-related non-random mating in a natural population of the ladybird beetle Harmonia axyridis (Coleoptera: Coccinellidae). Acta Societatis Zoologicae Bohemicae 79: 113-119.

Sawada Y., Aiba S., Takyu M., Repin R., Nais J., Kitayama K. (2015) Community dynamics over 14 years along gradients of geological substrate and topography in tropical montane forests on Mount Kinabalu, Borneo. Journal of Tropical Ecology 31:117?128.

Sawada Y., Aiba S., Seino T., Kitayama K.(2015) Size structure, growth and regeneration of tropical conifers along a soil gradient related to altitude and geological substrates on Mount Kinabalu, Borneo. Plan and Soil 10.1007/s11104-015-2722-z

Ushio M, Fujiki Y, Hidaka A, Kitayama K.(2015) Linkage of root physiology and morphology as an adaptation to soil phosphorus impoverishment in tropical montane forests. Functional Ecology 29: 1235?1245.
中西 麻美,稲垣 善之,柴田 昌三,大澤 直哉 (2015) ヒノキが優占する都市近郊二次林における小面積皆伐がリターフォールと堆積有機物に及ぼす影響. 森林応用研究 24: 1-9.


Hasegawa M, Ito M, Yoshida T, Seino T, Chung AYC, Kitayama k. (2014) The effects of reduced-impact logging practices on soil animal communities in the Deramakot Forest Reserve in Borneo. Applied Soil Ecology 83: 13-21

Iidzuka H., Goto H., Yamasaki M., Osawa N. (2014) Ambrosia beetles (Curculionidae: Scolytinae and Platypodinae) on Fagus crenata Blume: community structure, seasonal population trends and resource utilization patterns. Entomological Science 17: 167-180.

Imai N., Tanaka A., Samejima H., Sugau, J.B., Pereira J.T., Titin J., Kurniawan Y., Kitayama K. (2014) Tree community composition as an indicator in biodiversity monitoring of REDD+. Forest Ecology and Management 313: 169?179.

Kitayama K, Ando S, Repin R, Nais J. (2014) Vegetation and Climate of the Summit Zone of Mount Kinabalu in 1 Relation to the Walker Circulation. Arctic, Antarctic, and Alpine Research 46: 745-753.

Moles AT, Perkins SE, Laffan SW, Flores-Moreno H, Awasthy M, Tindall ML, Sack L, Pitman A, Kattge J, Aarssen LW, Anand M, Bahn M, Blonder B, Cavender-Bares J, Cornelissen JHC, Cornwell WK, Diaz S, Dickie JB, Freschet GT, Griffiths JG, Gutierrez AG, Hemmings FA, Hickler T, Hitchcock TD, Keighery M, Kleyer M, Kurokawa H, Leishman MR, Liu K, Niinemets U, Onipchenko V, Onoda Y, Penuelas J, Pillar VD, Reich PB, Shiodera S, Siefert A, Sosinski EE, Soudzilovskaia NA, Swaine EK, Swenson NG, van Bodegom PM, Warman L, Weiher E, Wright IJ, Zhang H, Zobel M, Bonser SP. 2014. Which is a better predictor of plant traits: Temperature or precipitation? Journal of Vegetation Science 25, 1167?1180.

Noriyuki S., Kameda Y., Osawa N. (2014) Male killer prevalence of the two sibling ladybird beetles Harmonia yedoensis and H. axyridis (Coleoptera: Coccinellidae) in the sympatric population. European Journal of Entomology 111: 307-311.

Onoda Y, Salunga JB, Akutsu K, Aiba S, Yahara T, Anten NPR. (2014) Tradeoff between light interception efficiency and light use efficiency? implication for species coexisting in one-sided light competition. Journal of Ecology 102: 167-175


Aiba S, Akutsu K and Onoda Y. (2013) Canopy structure of tropical and subtropical rainforests in relation to conifer dominance analysed with a portable LIDAR system. Annals of Botany 112: 1899-1909

Ando S., Kaneko S., Isagi Y., Repin R., Kitayama K. (2013) Development of SSR markers for the tropical alpine tree species Leptospermum recurvum (Myrtaceae) on Mount Kinabalu, Borneo. Applications in Plant Sciences 2013 1 ( 9 ): 1200010.

Aoyagi R., Nobuo Imai, Kanehiro Kitayama (2013), Ecological significance of the patched dominated by pioneer trees for the regeneration of dipterocarps in a Bornean logged-over secondary forest. Forest Ecology and Management 289, pp378-384.

Fujii S., Yamada A., Fukushima K, Saitoh S., Kitayama K., Takeda H. (2013) Relationships among distribution of fine roots, soil DOC concentration and Collembola. Plant Root

Hasegawa M, Ito M, Yoshida T, Seino T, Chung AYC, Kitayama K. (2013) The effects of reduced-impact logging practices on soil animal. Applied Soil Ecology

Hidaka A., Kitayama K. (2013) Relationship between photosynthetic phosphorus-use efficiency and foliar phosphorus fractions in tropical tree species. Ecology and Evolution (Open Access) doi: 10.1002/ece3.861

Kikuzawa K(*), Onoda Y(*), Wright IJ, Reich PB (2013) Mechanisms underlying global temperature-related patterns in leaf longevity. Global Ecology & Biogeography 28:982-993 (* equal contribution)

Kitayama K. (2013) The activities of soil and root acid phosphatase in the nine tropical rain forests that differ in phosphorus availability on Mount Kinabalu, Borneo. Plant and Soil

Kitayama K ed. (2013) Co-benefits of sustainable forestry ?Ecological studies of a certified Bornean rain forest- Springer, Tokyo.

Laugier G.J.M., Le Moguedec G., Tayeh A., Loiseau A., Osawa N., Estoup A., Facon B. (2013) Increase in male reproductive success and female reproductive investment in invasive populations of the harlequin ladybird Harmonia axyridis. Plos ONE 8: e77083. doi:10.1371/journal.pone.0077083.

Majuakim L., Kitayama K. (2013) Influence of polyphenols on soil nitrogen mineralization through the formation of bound protein in tropical montane forests of Mount Kinabalu, Borneo. Soil Biology & Biochemistry 57:14-21.

Pereira HM, Ferrier S, Walters M, Geller GN, Jongman RHG, Scholes RJ, Bruford MW, Brummitt N, Butchart SHM, Cardoso AC, Coops NC, Dulloo E, Faith DP, Freyhof J, Gregory RD, Heip C, Hoft R, Hurtt G, Jetz W, Karp DS, McGeoch MA, Obura D, Onoda Y, Pettorelli N, Reyers B, Sayre R, Scharlemann JPW, Stuart SN, Turak E, Walpole M, Wegmann M. Essential biodiversity variables. Science 339: 277-278.

Saori Fujii, Akinori Yamada, Keitaro Fukushima, Seikoh Saitoh, Kanehiro Kitayama and Hiroshi Takeda (2013) Relationships among distribution of fine roots, soil DOC concentration and Collembola. Plant Root

Suzuki S., Kitayama K., Aiba S-i., Takyu M., Kikuzawa K. (2013) Annual leaf loss caused by folivorous insects in tropical rain forests on Mt. Kinabalu, Borneo. Journal of Forest Research 18: 353-360.

Ushio M., Balser T., Kitayama K. (2013) Effects of condensed tannins in conifer leaves on the composition and activity of the soil microbial community in a tropical montane forest. Plant and Soil 365: 157-170

Yahara T, Javadi F, Onoda Y, de Queiroz LP, Faith DP, Prado DE, Akasaka M, Kadoya T, Ishihama F, Davies S. et al. (2013) Global legume diversity assessment: Concepts, key indicators, and strategies. Taxon 62: 249-266.


Aoki M., Fujii K., Kitayama K. (2012) Environmental control of root exudation of low-molecular weight organic acids in tropical rainforests. Ecosystems: 1-10.

Fujii K., Aoki M., Kitayama K. (2012) Biodegradation of low molecular weight organic acids in rhizosphere soils from a tropical montane rain forest. Soil Biology & Biochemistry 47: 142-148.

Gomyo M., Wakahara T., Shiraki K., Kuraji K., Kitayama K., Suzuki M. (2012) Comparison of nutrient balance between a tropical lowland forest and a tropical montane forest in Malaysian Borneo. Journal of Tropical Biology and Conservation 9: 15-25.

Imai N., Kitayama K., Titin J. (2012) Effects of logging on phosphorus pools in a tropical rainforest of Borneo. Journal of Tropical Forest Science 24: 5-17.

Imai N., Seino T., Aiba S-i., Takyu M., Titin J., Kitayama K. (2012) Effects of selective logging on tree species diversity and composition of Bornean tropical rain forests at different spatial scales. Plant Ecology: 1-12.

Kitayama K. (2012) Beta diversity of tree species along soil-P gradients in tropical montane rain forests of contrasting species pools: Does biodiversity matter in stabilizing forest ecosystems? Pacific Science 66: 151-160.

Langner A, Samejima H, Ong RC, Titin J, Kitayama K (2012) Integration of carbon conservation into sustainable forest management using high resolution satellite imagery:a case study in Sabah, Malaysian Borneo. International Journal of Applied Earth Observation and Geoinformation 18: 305-312.

Mizumachi E., Mori A.S., Akiyama R., Tokuchi N., Osawa N. (2012) Variation in herbivory-induced responses within successively flushing Quercus serrate seedlings under different nutrient conditions. Journal of Forest Research 17: 175-183.

Nicastro A, Onoda Y, Bishop MJ (2012) Direct and indirect effects of tidal elevation on eelgrass decomposition. Marine Ecology Progress Series456: 53-62.

Noriyuki S., Kawatsu K., Osawa N. (2012) Factors promoting maternal trophic egg provisioning in non-eusocial animals. Population Ecology 54: 455-465.

Noriyuki S., Osawa N. (2012) Intrinsic prey suitability in specialist and generalist Harmonia ladybirds: a test of the trade-off hypothesis for food specialization. Entomologia Experimentalis et Applicata 144: 279-285.

Noriyuki S., Osawa N., Nishida T. (2012) Asymmetric reproductive interference between specialist and generalist predatory ladybirds.Journal of Animal Ecology 81: 1077-1085.

Onoda Y, Lusk C. (2012) Like a jungle sometimes: how leaves survive in the rainforest understory. New Phytologist 195: 507-509

Onoda Y, Richards L, Westoby M. (2012) The importance of leaf cuticle for carbon economy and mechanical strength. New Phytologist 196: 441?447

Samejima H., Ong R., Lagan P., Kitayama K. (2012) Camera-trapping rates of mammals and birds in a Bornean tropical rainforest under sustainable forest management. Forest Ecology and Management 270: 248-256.

Suzuki S., Kitayama K., Aiba S-i., Takyu M., Kikuzawa K. (2012) Annual leaf loss caused by folivorous insects in tropical rain forests on Mt. Kinabalu, Borneo. Journal of Forest Research: 1-8.

Ushio M., Balser T.C., Kitayama K. (2012) Effects of condensed tannins in conifer leaves on the composition and activity of the soil microbial community in a tropical montane forest. Plant and Soil: 1-14.

Wagai R., Kitayama K. (2012) Soil organic matter storage in the tropical forests of Borneo: Pedogenic controls and ecological implications. Pedologist 55: 392-402.

Yamasaki M., Hiroaki Iizuka, Kazuyoshi Futai. (2012). Reproductive success of the ambrosia beetle Platypus quercivorus on Quercus laurifolia planted in Japan. Forest Research, Kyoto

de los Santos CB,Brun FG, Onoda Y, Cambridge ML, Bouma TJ, Vergara JJ, Perez-Llorens JL. (2012) Leaf-fracture properties correlated with nutritional traits in nine Australian seagrass species: potential implications for susceptibility to herbivory. Marine Ecology Progress Series. 458: 89?102

鈴木紀之、大澤直哉、西田隆義 (2012) 繁殖干渉による寄主特殊化の進化.日本生態学会誌 62: 267-274.